SOHO & Balance - DNS Settings Question

I am unclear about setting the LAN DNS. I have set the WAN DNS to OpenDNS . In this case, does every LAN/VLAN connection which is set to “automatic” will use whatever is specified for the WAN DNS? What is meant by the manual’s explanation: “The Pepwave router’s built-in DNS server address (i.e., LAN IP address) will be offered.” ?

Thank You!