The 8.5.1 release notes report issue 32917 resolved. For reference here is what is in the release notes:
32917: [System] Fixed an issue where the Local DNS Proxy used the hot standby WAN to resolve hostnames – All Models
Upgraded to 8.5.1 last Friday and today I checked usage on the backup cell connection which is in hot standby (standby priority 2). There was still over 40MB of usage per day even though the connection was never activated. I experimented and disabled DNS over HTTPS and now the traffic is completely gone from the cell backup link. This is on a B20x.
So I think issue 32917 has been resolved for normal DNS traffic, but is still a problem for DoH traffic, at least on the B20x. Please reopen the issue and get it resolved for the next release. Thank you.